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Psychology 337

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Title: Psychology 337


1
Psychology 337
  • Section 1 BackgroundA. Introduction

2
LANGUAGE
  • LANGUAGE FANCIEST CAPABILITY OF LIVING FORMS
  • MAIN OBVIOUS DEFINING BEHAVIOR OF HUMANS WHO ARE
    THE MOST ADVANCED SPECIES
  • ALLOWS UNIVERSAL COMMUNICATION OF MEANINGS
  • ANOTHER NEW HUMAN STEP MATERIAL CULTURE HUGE
    SIGNIFICANCE
  • MOST RECENTLY (5,000 YA) WRITTEN LANGUAGE

3
LANGUAGE PLUS CULTURE
  • GIVE STUPENDOUS INTERGENERATIONAL KNOWLEDGE
    TRANSMISSION
  • A NEW FORM OF EVOLUTION WITH COMPOUND INTEREST
  • WE STAND ON THE SHOULDERS OF GIANTS (NEWTON)

4
PSYCHOLINGUISTICS
  • SCIENTIFIC STUDY NEW (1962) AND PRELIMINARY
  • PSYCHOLINGUISTICS DEFINITION THE SCIENCE OF
    HUMAN LANGUAGE PRODUCTION, COMPREHENSION AND
    ACQUISITION (LEVELT, 1989)
  • SPEECH - LANGUAGE DISTINCTION SPEECH IS THE
    NORMAL PRIMARY MEDIUM OF LANGUAGE. (NOTE ALSO
    SIGN LANGUAGE OF THE DEAF) READING AND WRITING
    CONSTITUTE A SECONDARY (TAUGHT) MEDIUM

5
  • THE COURSE IS ORGANIZED AROUND A FUNDAMENTAL
    DIFFERENCE OF OPINION BETWEEN TWO SCHOOLS OF
    THOUGHT 1.FORMALISM AND 2.NEODARWINIANISM
  • 1.FORMALISM
  • ALTERNATELY
  • ESSENTIALISM, RATIONALISM, OBJECTIVISM. LONG
    HISTORY A.PLATO, B.DESCARTES, AND C. CHOMSKY
  • THESISTHE FORM OF THINGS IS A'PRIORI

6
  • A) PLATO THINGS HAVE ESSENCES (TRIANGLE)
  • B) DESCARTES ALL KNOWLEGE AMENABLE TO
    MATHEMATICALLY DRIVEN REASON (OF HUMANS)
  • C) CHOMSKY LANGUAGE RESULTS FROM INNATE
  • UNIVERSAL GRAMMAR

7
  • 2.NEODARWINIANISM IS A MODERN CONSENSUS ON
    DARWIN'S THEORY OF EVOLUTION BY NATURAL
    SELECTION.
  • THESIS FORM OF ALL LIVING THINGS ARISES FROM
    SUCCESSFUL FUNCTION (SURVIVAL OF THE FITTEST)

8
THE 2 SCHOOLS OF THOUGHT
1. FORMALISM LANGUAGE GENETICALLY SPECIFIED
EXPERIENCE TRIVIAL.
2. NEODARWINIANISM INTERACTION WITH WORLD
CRUCIAL
(INCLUDING CULTURE).
Environment
Environment
Organism
Organism
9
  • THESE TWO VIEWS ARE INCOMPATIBLE, AND IN THE
    ABSENCE OF OTHER ALTERNATIVES, ONE OF THEM IS
    PROBABLY RIGHT.
  • I THINK DARWIN WAS RIGHT
  • THE FORCE OF THE FORMALIST ARGUMENT ABOUT
    LANGUAGE COMES MAINLY FROM POINTING TO LINGUISTIC
    PHENOMENA, LARGELY IN LANGUAGE ACQUISITION, THAT
    AREN'T PRESENTLY UNDERSTANDABLE IN ANY OTHER WAY
    THAN INVOKING A'PRIORI FORM.

10
BEWARE HELMONTS ERROR
  • BUT NOT UNDERSTANDABLE DOESNT MEAN IT MUST BE
    APRIORI
  • CONSIDER HELMONTS ERROR
  • HE DID AN EXPERIMENT THAT SHOWED THAT TREES
    GROW BY WATER ALONE
  • HE LEFT OUT 2 CRUCIAL BUT NON-OBVIOUS
    VARIABLES
  • SUNLIGHT AND AIR

11
SELF ORGANIZATION
  • ANOTHER POSSIBILITY FOR THE CREATION OF FORM.
  • DEFINITION A SELF ORGANIZING SYSTEM IS ONE IN
    WHICH SOME KIND OF HIGHER ORDER PATTERN EMERGES
    FROM THE INTERACTIONS OF MULTIPLE SIMPLE
    COMPONENTS WITHOUT THE BENEFIT OF A LEADER,
    CONTROLLER OR ORCHESTRATOR. (CLARK, 1997)
  • SELF-ORGANIZATION IS FOUND IN PHYSICAL AND
    BIOLOGICAL WORLDS

12
EXAMPLES OF SELF-ORGANIZATION
  • EXAMPLE 1 HEXAGONAL CELLS IN BEEHIVES
  • (SEE READING ON THE CAUSES OF FORM IN NATURE)
  • EXAMPLE 2 ELEVATOR PRINCIPLE OF VOWEL SYSTEMS

13
MY CONCLUSION
  • AS EVERY OTHER MAJOR ASPECT OF LIFE FORMS EVOLVED
    BY NATURAL SELECTION, LANGUAGE PROBABLY DID TOO.
  • BUT WHOEVER IS RIGHT, THE CONTEST BETWEEN THESE
    APPROACHES IS A MAJOR ASPECT OF CURRENT COGNITIVE
    SCIENCE.

14
BRAIN-LANGUAGE FOCUS
  • CROSS-SPECIES COMPARISON OF BRAIN EVOLUTION
    TOGETHER WITH BEHAVIORAL EVOLUTION GIVES A MORE
    COMPLETE EVOLUTIONARY PICTURE.
  • HELPS COMPENSATE FOR THE FACT THAT LANGUAGE
    DOESN'T FOSSILIZE.
  • OTHER PRIMATES (PROSIMIANS, MONKEYS AND APES) CAN
    BE TAKEN TO BE SIMILAR TO OUR ANCESTORS.

15
BASIC PROPOSITION OF THIS COURSE
  • HUMAN LANGUAGE IS UNIQUE, BUT NEVERTHELESS MUST
    HAVE EVOLVED FROM NONLINGUISTIC CAPABILITIES
    (A'PRIORI FORM IS ALSO UNEXPLAINABLE FORM)
  • FIGURING OUT HOW WILL ULTIMATELY LEAD TO THE BEST
    UNDERSTANDING OF PSYCHOLINGUISTICS. SO WE SHOULD
    TRY TO DO IT
  • THIS IS THE "ULTIMATE CAUSES" PERSPECTIVE OF MAYR

16
Psychology 337
  • Section 1 BackgroundB. Language Definition

17
LANGUAGE DEFINITION
  • DEFINITION (SPOKEN) FEWER THAN 150 SOUNDS
    (CONSONANTS AND VOWELS MODE 30. RANGE 11-141)
    WHICH ARE INDIVIDUALLY MEANINGLESS, ARE
    CONCATENATED (PUT TOGETHER), NOT IN ALL POSSIBLE
    WAYS, TO PRODUCE SOME HUNDREDS OF THOUSANDS OF
    MEANINGFUL MORPHEMES (MEANING UNITS WORDS OR
    PARTS OF WORDS) WHICH HAVE MEANINGS THAT ARE
    ARBITRARILY ASSIGNED, AND THEY ARE CONCATENATED
    BY RULE TO YIELD AN INFINITE SET OF SENTENCES,
    HAVING MEANINGS THAT CAN BE DERIVED."
  • (ROGER BROWN - WITH MODIFICATIONS)

18
LANGUAGE4 IMPORTANT PROPERTIES
  • 1. ARBITRARINESS LACK OF A PRINCIPLED RELATION
    BETWEEN A CONCEPT AND THE SOUND PATTERN THAT GOES
    WITH IT.
  • ENGLISH "CHURCH" FRENCH "EGLISE" ETC
  • CONTRAST THIS WITH ICONICITY SIGNAL RESEMBLE
    CONCEPT. E.G. SOME SIGNS IN SIGN LANGUAGE
  • HISTORY OF THOUGHT ON THIS PROBLEM
  • PREDARWINIAN IN "ADAMIC LANGUAGE" THERE WAS A
    PRINCIPLED RELATION BETWEEN CONCEPTS AND SOUND
    PATTERNS. THEN CAME THE TOWER OF BABEL.
  • DARWIN FORMS MUST ARISE FROM USE

19
  • LANGUAGE IS NOT ENTIRELY ARBITRARY
  • AND IT COULDNT HAVE STARTED ARBITRARY
  • 1. ONOMATOPOEIA
  • 2. PHONETIC SYMBOLISM LOW FREQUENCY SOUNDS FOR
    BIG OBJECTS AND VICE VERSA
  • 3. "MAMA" CONTRIBUTED BY BABIES
  • BUT THIS DOESN'T ACCOUNT FOR MUCH

20
  • THERE MUST HAVE BEEN A LOT OF INVENTION OF WORDS
    BY CHIEFS, WITCH DOCTORS ETC
  • ALSO METAPHOR MUST HAVE PLAYED EVENTUALLY PLAYED
    A HUGE ROLE. E.G. CONCRETE TO ABSTRACT
  • MY THEORY THE INITIAL RAW MATERIAL FOR WORDS WAS
    LIKE THE BABBLING OF BABIES
  • IN FACT THE VERY FIRST WORDS MIGHT HAVE BEEN
    INVENTED IN A BABY TALK SITUATION
  • ONE IMPORTANT IMPLICATION OF ARBITRARINESS
  • INFANTS MUST LEARN THE WORDS OF LANGUAGE X
    ONE-BY-ONE - THE LINKAGES CANT BE INNATE

21
  • 2. DISPLACEMENT
  • THE SUBJECT MATTER OF LANGUAGE IS NOT RESTRICTED
    TO THE HERE AND NOW
  • WE CAN TALK ABOUT OTHER PLACES AND TIMES, AND
    EVEN ABOUT THE NON-REAL
  • THIS SETS LANGUAGE APART FROM OTHER OTHER FORMS
    OF ANIMAL COMMUNICATION

22
  • 3. DUALITY TWO LEVELS OF ORGANIZATION
  • A. MEANINGLESS SOUNDS INTO MEANINGFUL MORPHEMES
    (PHONOLOGY)
  • B. MORPHEMES INTO SENTENCES

23
  • 4. PRODUCTIVITY CONCATENATION AT EACH OF THE TWO
    LEVELS, EACH WITH RULES OF COMBINATION.
    (COMBINATION AN ORDERED SEQUENCE)
  • EXAMPLES IN ENGLISH
  • PHONOLOGY (3 SOUNDS - 3 POSSIBILITIES)
  • "CAT", "TACK", "ACT"
  • SYNTAX "JOHN HIT MARY" "MARY HIT JOHN" OR
    "BLIND VENETIAN" "VENETIAN BLIND"

24
  • MORE ON PRODUCTIVITY
  • NOTE THE POWER OF COMBINATION TO "PRODUCE (OR
    GENERATE) LARGE NUMBERS OF ENTITIES.
  • THE NUMBER OF POSSIBLE COMBINATIONS OF N UNITS
    USED ONCE IN EACH SET IS N! (N FACTORIAL)
    EXAMPLES FOLLOW.
  • 3 UNITS A B C
  • COMBINATIONS (6) ABC ACB BAC BCA CAB CBA
  • 3! 3 X 2 X 1 6
  • 4! 4 X 3 X 2 X 1 24.
  • 10! 10 X 9 X 8 ... 3.628.800 ETC

25
  • NOW IMAGINE THE POSSIBILITIES IF YOU COULD USE
    EACH UNIT MORE THAN ONCE, IN SETS OF VARIOUS
    SIZES, NOT JUST SIZE 3 AS IN THE ABOVE EXAMPLE.
  • THUS, HUNDREDS OF THOUSANDS OF MORPHEMES ARE
    POSSIBLE.
  • AN INFINITE NUMBER OF SENTENCES IS MADE
    MATHEMATICALLY POSSIBLE BY MEANS OF RECURSION THE
    SAME STRUCTURE CAN RECUR IN A SINGLE SENTENCE.
    TAKE THE FOLLOWING EXAMPLE

26
  • 1.This is the house that Jack built.
  • 2.This is the mouse that lived in the house that
    Jack built.
  • 3.This is the rat that worried the mouse that
    lived in the house that Jack built."
  • 4.This is the cat that ...etc.
  • WITH EACH NEW "THAT" WHICH OCCURS SUBSEQUENT TO
    SENTENCE 1, A RELATIVE CLAUSE RECURS.
  • THOUGH WE DON'T USUALLY DO MANY RELATIVE CLAUSES
    AT A TIME IN ONE SENTENCE, ITS EASY TO SEE, FROM
    THE PROPERTIES OF COMBINATION THAT WE CAN PRODUCE
    AN INDEFINITE NUMBER OF SENTENCES, AND WHY
    HOCKETT SAID THAT MOST SENTENCES THAT ARE UTTERED
    WERE NEVER UTTERED BEFORE AND WILL NEVER BE
    UTTERED AGAIN.

27
  • NOTE THE CONSEQUENCES OF THIS FOR OUR
    UNDERSTANDING OF LANGUAGE ACQUISITION.
  • INFANTS COULDN'T LEARN EACH OF THEIR INDIVIDUAL
    SENTENCES (E.G. BY IMITATION). THEY MUST LEARN
    RULES FOR SENTENCE CONSTRUCTION.
  • AN ANALOGY HAS BEEN MADE BETWEEN LANGUAGE
    STRUCTURE AND GENETIC STRUCTURE.
  • GENES (100,000 IN HUMANS) ARE MADE UP OF DNA
    WHICH CONSISTES BASICALLY OF UNLIMITED
    COMBINATIONS OF JUST FOUR COMPOUNDS ADENOSINE,
    THYMIDINE, CYTOSINE AND GUANINE.
  • THERE IS ALSO AN ANALOGY WITH THE ELEMENTS OF
    CHEMISTRY.

28
THE PARTICULATE PRINCIPLE
  • THE PRINCIPLE BY WHICH UNITS ARE CONCATENATED
    INTO FORMS THAT HAVE PROPERTIES OF THEIR OWN,
    OVRE AND ABOVE THE PROPERTIES OF THE UNITS THAT
    COMPOSE THEM IS KNOWN AS THE PARTICULATE PRINCIPLE

29
Psychology 337
  • Section 1 BackgroundC. Evolution

30
COMPARISON OF LANGUAGE WITH VOCAL COMMUNICATION
IN OTHER PRIMATES
  • OTHER PRIMATES HAVE A REPERTOIRE OF ABOUT 30
    CALLS WITH ONE CALL PER MEANING UNDER NATURAL
    CONDITIONS (WE WILL DEAL WITH ATTEMPTS TO TEACH
    OTHER PRIMATES LANGUAGE LATER).
  • THUS, OTHER PRIMATES HAVE NEITHER DUALITY NOR
    PRODUCTIVITY. LITTLE DISPLACEMENT EITHER.

31
  • UNLIKE HUMANS, EACH OF WHOM LEARNS THE LANGUAGE
    OF THEIR PARTICULAR COMMUNITY, OTHER PRIMATES,
    SEEM TO HAVE NEGLIGIBLE VOCAL LEARNING CAPACITY.
  • (THIS IS SHOWN BY CROSS-FOSTERING MONKEY INFANTS
    OF DIFFERENT SPECIES THE INFANTS DO NOT LEARN
    THE CALLS OF THEIR FOSTER PARENTS.)
  • THUS TO ME, THE MAIN PROBLEM IN THE EVOLUTION OF
    LANGUAGE IS TO EXPLAIN HOW WE EVOLVED FROM
    ANIMALS THAT DID NOT HAVE DUALITY, PRODUCTIVITY
    OR DISPLACEMENT IN THEIR COMMUNICATION SYSTEM TO
    ANIMALS THAT DO, ACQUIRING A GREATER LEARNING
    CAPACITY AND MORE INDEPENDENCE OF COMMUNICATIVE
    SIGNALS FROM EMOTIONAL STATES IN THE PROCESS.

32
WHAT DO ANIMALS HAVE?
  • BUT NOTICE THAT MANY OTHER ANIMALS HAVE
    NONLINGUISTIC COGNITIVE CAPACITIES THAT POWER
    LANGUAGE IN HUMANS.
  • TO TAKE SOMETHING THAT OCCURS IN HUMAN AND OTHER
    PRIMATE ENVIRONMENTS, FECES IS SOMETHING THAT ALL
    PRIMATES HAVE A CONCEPT OF, OR A SPECIFIC
    CATEGORY FOR. WORDS MUST ATTACH TO SUCH
    NONLINGUISTIC CONCEPTS OR CATEGORIES TO HAVE
    MEANING.

33
  • SECOND, LINGUISTIC EVENTS ARE TYPICALLY TAILORED
    TO THEIR SOCIAL ENVIRONMENT ("PRAGMATICS" IS THE
    STUDY OF THIS) BUT BEHAVIOR IN GENERAL, IN
    HUMANS, AND OTHER PRIMATES IS TYPICALLY TAILORED
    TO THE SOCIAL ENVIRONMENT.
  • THUS, LANGUAGE MUST INTERFACE WITH NONLANGUAGE IN
    TWO MAIN WAYS, AND WE MUST STUDY HOW.

34
EVOLUTION
  • DARWIN'S THEORY OF EVOLUTION BY NATURAL SELECTION
    IS AS POWERFUL AND IMPORTANT AS ANY THEORY IN THE
    HISTORY OF SCIENCE (E.G. NEWTON'S EINSTEIN'S).
  • IT PROVIDES A POSSIBLE EXPLANATION (EITHER
    ALREADY AVAILABLE OR IN PRINCIPLE) OF THE ORIGIN
    AND NATURE OF ALL LIVING FORMS. (THEREFORE - IN
    PRINCIPLE - FOR LANGUAGE).

35
OPINIONS ON THE THEORY
  • 1. "HISTORICAL PERIODS ARE DOMINATED BY DISTINCT
    SETS OF IDEAS. GREEK PHILOSOPHY, CHRISTIANITY,
    RENAISSANCE THOUGHT, THE SCIENTIFIC REVOLUTION,
    AND THE ENLIGHTENMENT ARE SETS OF IDEAS THAT
    DOMINATED THEIR HISTORICAL PERIOD. ... THE
    CHANGES ... ARE OFTEN REFERRED TO AS REVOLUTIONS.
    THE MOST FAR REACHING OF ALL THESE INTELLECTUAL
    UPHEAVALS WAS THE DARWINIAN REVOLUTION."
  • ERNST MAYR - MOST EMINENT LIVING EVOLUTIONARY
    THEORIST

36
  • 2. ON DARWIN - "A BRILLIANT MIND, GREAT
    INTELLECTUAL BOLDNESS, AND AN ABILITY TO COMBINE
    THE BEST QUALITIES OF A NATURALIST-OBSERVER,
    PHILOSOPHICAL THEORETICIAN, AND EXPERIMENTALIST -
    THE WORLD HAS SO FAR SEEN SUCH A COMBINATION ONLY
    ONCE, AND IT WAS IN THE MAN CHARLES DARWIN.
  • MAYR
  • 3. DARWIN'S THEORY -"OUTLINED THE ESSENTIAL
    FEATURE OF THE SECRET OF LIFE."
  • FRANCIS CRICK - NOBEL PRIZE WINNER FOR
    CO-DISCOVERY OF DNA STRUCTURE

37
  • 4. "THIS BOOK IS WRITTEN IN THE CONVICTION THAT
    OUR OWN EXISTENCE ONCE PRESENTED THE GREATEST OF
    ALL MYSTERIES, BUT THAT IT IS A MYSTERY NO LONGER
    BECAUSE IT IS SOLVED. DARWIN AND WALLACE SOLVED
    IT ... I WROTE THIS BOOK BECAUSE I WAS SURPRISED
    THAT SO MANY PEOPLE SEEMED NOT ONLY UNAWARE OF
    THE ELEGANT AND BEAUTIFUL SOLUTION TO THIS
    DEEPEST OF PROBLEMS BUT, INCREDIBLY, IN MANY
    CASES ACTUALLY UNAWARE THAT THERE WAS A PROBLEM
    IN THE FIRST PLACE.
  • RICHARD DAWKINS - AUTHOR OF THE BEST MODERN
    DEFENSE OF THE THEORY
  • 5. "IF I WERE TO GIVE AN AWARD FOR THE SINGLE
    BEST IDEA ANYONE HAS EVER HAD, I'D GIVE IT TO
    DARWIN, AHEAD OF NEWTON, EINSTEIN, AND EVERYONE
    ELSE". DENNETT, 1995, P 21.

38
PRESENT STATUS OF THE THEORY
  • IN SPITE OF ITS MERITS IT IS OFTEN IGNORED,
    DISPUTED AND MISUNDERSTOOD.
  • TWO REASONS FOR THIS
  • 1. IT CONTRADICTS RELIGIOUS IDEAS. GALLUP 50
    OF AMERICANS (25 OF COLLEGE GRADUATES) BELIEVE
    THE CREATION SCENARIO ("1 NATION UNDER GOD").
  • 2. IT OPPOSES CONCEIT. MOST HUMANS TEND TO BE
    ANTHROPOCENTRIC - THEY WANT TO BELIEVE THEY ARE A
    CLASS APART FROM ANY OTHER LIFE FORM, (AND
    DESCARTES HAS ENCOURAGED THEM).

39
  • THERE CERTAINLY SEEM TO BE GOOD REASONS TO
    BELIEVE WE ARE DIFFERENT.
  • BUT THE QUESTION IS HOW DID WE GET DIFFERENT.
  • EVOLUTION IS A FACT AS WELL DOCUMENTED AS
    ANYTHING ELSE WE CALL A FACT.
  • DENNETT ANYONE WHO DOESN'T KNOW THIS IS SIMPLY
    IGNORANT.

40
TWO FACTS MAKE EVOLUTION A FACT
  • 1. FOSSIL RECORD SIMPLER FORMS IN LOWER
    STRATA MORE COMPLEX BUT OBVIOUSLY RELATED FORMS
    IN HIGHER STRATA.
  • 2. RADIOACTIVE DATING TECHNIQUES SHOW AGE OF
    STRATA - LOWER STRATA EARLIER.
  • THEREFORE MORE COMPLEX FORMS EVOLVED FROM MORE
    SIMPLE FORMS IN THE TIME DOMAIN.

41
DARWINS TWO CONTRIBUTIONS
  • 1. HE PRESENTED MOUNTAINS OF EVIDENCE FOR THE
    FACT OF EVOLUTION
  • THIS FIRST CONTRIBUTION HAS BEEN BASICALLY
    ACCEPTED IN THE SCIENTIFIC COMMUNITY ALMOST FROM
    THE TIME HE MADE IT (1859 "THE ORIGIN OF
    SPECIES".)
  • 2. HE SUGGESTED A MECHANISM FOR EVOLUTION -
    NATURAL SELECTION.
  • THIS HAS REMAINED CONTROVERSIAL BUT IS NOW FOR
    THE MOST PART ACCEPTED, AND THERE IS NO VIABLE
    ALTERNATIVE.

42
WHAT IS NATURAL SELECTION?
  • SHORTHAND "SURVIVAL OF THE FITTEST" (SPENCER).
  • MEMBERS OF A SPECIES EXHIBIT VARIATION IN MANY
    ATTRIBUTES (E.G HEIGHT) (DISTRIBUTION OF
    VARIATION NORMAL CURVE).
  • MEMBERS IN A CERTAIN SUB-RANGE OF THE RANGE OF
    VARIATION MAY BE BETTER ABLE TO RESPOND TO
    ENVIRONMENTAL PRESSURES THAN OTHERS, AND
    CONSEQUENTLY REPRODUCE MORE OFFSPRING THAN
    OTHERS. OVER A NUMBER OF GENERATIONS THIS TREND
    WILL LEAD TO NEW SPECIES.

43
EXAMPLES
  • 1. TALLER GIRAFFES CAN FEED OFF LEAVES HIGHER
    IN TREES.
  • 2. MOTHS CHANGE COLOR TO MATCH TREES AS
    POLLUTION LEVELS CHANGE (CAMOUFLAGE).
  • 3. FOLLOWING FOOD SHORTAGES ON THE GALAPAGOS
    ISLANDS, SPECIES OF DARWIN'S FINCH WITH BEAK
    CHARACTERISTICS BEST SUITED TO EXPLOITING
    REMAINING FOOD HAD RELATIVELY MORE OFFSPRING IN
    THE NEXT GENERATION (SEE GRANT) THIS IS ON-LINE
    EVOLUTION.

44
(No Transcript)
45
KEY TERMS
  • 1.ENVIRONMENTAL PRESSURES "SELECTION PRESSURES"
  • 2. SUCCESSFUL CHANGES "ADAPTATIONS"

46
VARIATION (ESSENTIAL)
  • 2 SOURCES
  • 1. MUTATIONS (CHANGES IN GENES) MOST ARE FATAL.
    LARGE CHANGES IMPOSSIBLE BECAUSE CHANGES MUST BE
    COMPATIBLE WITH EXISTING PROPERTIES FOR THE
    RESULT TO WORK.
  • 2. SEXUAL REPRODUCTION. (MORE IMPORTANT) GIVES
    NEW MIXES OF GENES.

47
The Origin of Species(The modern theory of
biological evolution)
Propagation
Genetic Processes
Variation
Natural Selection
Species
Adaptations
E.g. Mutation, Sex
Selective Pressure
Anatomy Behavior
48
THREE PROBLEMS IN UNDERSTANDING THE THEORY
(CRICK)
  • 1. HARD TO CONCEIVE THE SLOWNESS OF THE TIME
    SCALE - BIG EFFECTS BY SMALL STEPS. (NOTE HOYLE'S
    PROBLEM OF BOEING 707S FROM JUNKYARDS) BUT,
    SIZEWISE, FROM A MOUSE TO AN ELEPHANT BY 1 1/2
    OZ. STEPS WOULD TAKE 12.000 5 YEAR GENERATIONS
    -HARDLY ANY TIME IN EVOLUTIONARY TERMS.

49
  • 2. Q HOW CAN RANDOM VARIATION GIVE COMPLEX
    STRUCTURE? A. VARIATION IS RANDOM BUT SELECTION
    IS NOT.
  • 3. PROBLEM HOW COULD EACH STEP TOWARDS AN "ORGAN
    OF EXTREME COMPLICATION AND PERFECTION" (E.G. THE
    EYE) BE ADAPTIVE? 1 VISION IS BETTER THAN NO
    VISION.

50
3.8
EVOLUTION OF LIFE FORMS (LEWIN, CH 1)
51
PRIMATE EVOLUTION (LEWIN, CH 4)
52
Prosimians
Aye aye
Red-fronted lemur
Tarsier
Slow loris
Sifaka
Ring-tailed lemur
53
Gibbons Siamangs
Pigmy marmoset
Golden-lion tamarin
Chimpanzee
Squirrel monkey
Baboon
Orangutan
Mountain gorilla
Spider monkey
Japanese macaque
Colobus monkey
Rhesus macaque
54
(possibly 1.2 mya)
HUMAN EVOLUTION (LEWIN, CH 7)
55
The Hominid Family Tree
  • Question When did the hominid family tree split
    from a common ancestor with chimpanzees?
  • DNA dating estimate 5-7 mya.
  • Recent fossil evidence points more to 7 mya.

56
Bipedalism
  • Bipedalism with upright posture is considered to
    be the main initial defining property of humans.
  • It has been considered to have been selected to
    allow early hominids to see better on the
    savanna, to facilitate long journeys, to give
    more protection of the body from the sun, and to
    allow hominids to carry food and weapons.
  • All but the last of these has been ruled out by
    recent findings suggesting bipedalism in hominids
    living in the forest close to 5 mya.

57
Bipedalism and I.Q.
  • Another surprise has been that while logic
    suggests the bipedalism must have been associated
    with an increase in the brain-size to body-size
    ratio--the Encephalization Quotient or E.Q--brain
    size increases didnt begin before Homo Habilis
    at about 2 mya.

58
Lumpers vs Splitters
  • Paleontologists can be divided in lumpers,
    postulating as few forms as possible, and
    splitters, postulating more forms.
  • Recent findings suggest more forms than before.
    But there is some evidence of a major split into
    2 lines The Robust line with animals with huge
    prognathous jaws and small brains, and the
    Gracile line with smaller less protruding jaws,
    beginning with Homo Habilis.
  • It is thought that the evolution of tool use took
    the burden of food processing off the jaws, but
    the earliest tools have been found at Robust
    sites.

59
Out of Africa
  • Recent evidence suggests that there have been a
    number of migrations out of Africa, starting
    close to 2 mya and ending with the last one at
    only about 80 tya. The main homo line in Africa
    at about 2 mya is now called Ergaster rather than
    Erectus.
  • The term Erectus is reserved for the migrating
    populations, up to about 300 tya, when modern
    Homo Sapiens - our species - first arose.

60
Tool Use
  • Tool use was taken to be a major defining
    property of humans, but many other animals are
    now known to use tools.
  • Jane Goodalls chimpanzees, who take leaves off
    twigs before thrusting them into termite mounds
    to get termites, also manufacture tools. Now the
    only defining human characteristic relative to
    tools is that only we make machine tools (tools
    to make tools).

61
Brain Size
  • The rate of increase in brain size in the last 2
    mya (unprecedented in other life forms) is
    presently not well understood. It cant be tied
    to developments in tool manufacture and use,
    because they go in fits and starts.
  • One suggestion Its related to dietary
    improvements involving tubers.
  • Another suggestion Its related to
    survival-related increases in social
    intelligence. E.g Theory of Mind.
  • Most recent suggesting It is directly produced
    by genetic changes.

62
Use of Fire
  • The use of fire, now dated to 1.2 mya (Brain) was
    an important development for protection from
    predation, food preparation and nutrition, and
    the probable socio-cultural developments that
    resulted from groups sitting around camp fires
    for long periods of time.

63
When Language Evolved
  • Two extreme lines of thought
  • 1. Paleontologists (stone and bone people, e.g.
    Holloway) think it might have evolved in Homo
    Habilis, because there is endocast evidence for a
    Brocas area in this taxon.
  • 2. Anthropologists (E.G. Klein) think language
    might have evolved in the last 50 tya, because it
    must have accompanied the apparent cultural
    explosion occurring at about this time, judging
    from new tools, cave drawings, figurines, burial
    of the dead with bodily ornaments etc (e.g. at
    Lascaux in southern France and Altamira in
    northern Spain).

64
Mono- or Polygenesis of Language
  • There is some consensus that there might have
    been a single first language evolving in Africa
    before 80 tya, and subsequently spreading to the
    rest of the world.

65
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ENCEPHALIZATION QUOTIENT (DONALD, CH 4)
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EVOLUTION OF COMMUNICATION USE OF MEDIA
  • OLFACTORY- IMPORTANT IN NOCTURNAL MAMMALS
  • VISUAL
  • 1. POSTURAL - BASIC IN TETRAPODS
  • 2. FACIAL- DEVELOPSIN MAMMALS IMPORTANTINTERRESTRI
    AL PRIMATES
  • 3. GESTURAL (MANUAL) ONLY IMPORTANT IN HUMANS
  • AUDITORY DEVELOPS IN MAMMALS (ISOLATION PEEP)
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