Title: Ranunculaceae Buttercup Family
1Ranunculaceae (Buttercup Family)
Habit herbs, shrubs, or occational vines Stems
with vascular bundles in concentric rings Leaves
usually alternate and spiral, simple, sometimes
lobed or dissected, to compound Inflorescence
determinate Flowers usually bisexual, radial to
occationally bilateral, with short to elongate
receptacle Perianth parts usually not 3-merous,
tepals 4 to numerous, distinct or perianth
differentiated into calyx and corolla, then
sepals usually 5, distinct and deciduous, and
petals usually 5, distinct, often with nectar
producing basal portion Stamens numerous,
anthers opening by longitudinal slits Pollen
tricolpate Carpels usually 5 to numerous,
occationally reduced to 1, usually distinct,
ovaries superior, with usually lateral
placentation
2Fruit usually an aggregate of follicles or
achenes. Floral Formula Floral Diagram
3Distribution and Ecology Widespread, but
especially characteristic of temperate and boreal
regions of the Northern Hemisphere. Genera/specie
s 47/2000 Major genera Ranunculus (400 spp.),
Aconitum (250 spp.), Clematis (250 spp.),
Delphinium (250 spp.), Anemone (150 spp.), and
Thalictrum (100 spp.). Economic plants and
products The family is chiefly important for its
numerous ornamental herbs, such as Anemone
(windflower), Aconitum (monkshood), Actaea
(baneberry), Aquilegia (columbine), Caltha (marsh
marigold), Clematis (virgins bower), Delphinium
(larkspur), Helleborus (hellebore), Ranunculus
(buttercup), Thalictrum (meadow rue), and
Trollius (globeflower). A number of genera are
highly poisonous.
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6Ranunculus ficaria
7Ranunculus septentrionalis
8Isopyrum biternatum
9Isopyrum biternatum
10Eranthis hymalis
11Eranthis hymalis
12Coptis groenlandica
13Caltha palustris
14Caltha palustris
15Anemone canadensis
16Anemone cylindrica
17Anemone cylindrica
18Anemone americana
19Anemone sp.
Anemone nobilis
20Clematis reticulatis
Clematis virginiana
21Helleborus purpurascens
Hydrastis canadensis
22Aconitum cultivar
23Aquilegia canadensis
24Aquilegia canadensis
25Aquilegia canadensis
26Delphinium ajacis
27Delphinium Streichen
28Delphinium Streichen
29Thalictrum revolutum
30Thalictrum revolutum
31Thalictrum basicarpum
32Papaveraceae (Poppy Family)
Habit herbs to soft-wooded shrubs Stems with
vascular bundles sometimes in several rings,
with laticifers present and plants with white,
cream, yellow, orange, or red sap, or with
specialized elongated secretory cells and sap
then mucilaginous, clear Leaves usually
alternate and spiral, simple, but often lobed or
dissected, sometimes spinose Inflorescence
various Flowers bisexual, radial to
bilateral Sepals usually 2 or 3, usually quickly
deciduous Petals usually 4 or 6, sometimes
numerous, distinct, often crumpled in bud and
thus wrinkled when expanded, often the 2 (or 3)
inner differentiated from the 2 (or 3) outer
33Stamens numerous, to 6 that are connate in 2
groups of 3 Pollen tricolporate to polyporate
Carpels 2 to numerous, connate, ovary superior,
with parietal placentation Fruit a capsule,
opening variously, but often with apical pores,
valves, or longitudinal slits, often with a
persistent thickened replum, seeds somewhat
arillate. Floral Formula
34Floral Diagram Distribution and
Ecology Widely distributed in mainly temperate
regions, especially diverse in the Northern
Hemisphere, but also in southern Africa and
eastern Australia. Genera/species 40/770
35Major genera Corydalis (400 spp.), Papaver (100
spp.), Fumaria (50 spp.), and Argemone (30 spp.).
Genera occurring in the continental United
States and/or Canada include Argemone, Corydalis,
Dendromecon, Dicentra, Eschscholzia, Fumaria,
Papaver, and Romneya, among others. Delimitation
of genera is often difficult. Economic plants
and products Papaver somniferum (opium poppy) is
the source of opium and derivatives such as
morphine, heroin, and codeine the seeds of this
species (which do not contain opium) are used as
a spice. Many have showy flowers and are
cultivated as ornamentals, such as species of
Argemone (prickly poppy), Eschscholzia
(California poppy), Papaver (poppy), Macleaya
(plume poppy), Corydalis (harlequin), Sanguinaria
(bloodroot), and Dicentra (Dutchmans-breeches).
Most species are highly poisonous.
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38Argemone albiflora
39Argemone albiflora
40Chelidonium majus
41Stylophorum diphyllum
42Sanguinaria canadensis
43Sanguinaria canadensis
44Eschscholtzia caespitosa
Papaver nudicaule
45Papaver orientale
46Dicentra canadensis
Dicentra specabilis
47Dicentra canadensis
48Corydalis micrantha
49Caryophyllaceae (Carnation or Pink Family)
Habit usually herbs Stems sometimes with
concentric rings of xylem and phloem,
anthocyanins present Leaves opposite, simple,
entire, often narrow, the leaf pair often
connected by a transverse nodal line, and nodes
usually swollen Inflorescence determinate,
terminal Flowers usually bisexual, radial,
sometimes with an androgynophore Tepals 4-5,
distinct to connate, usually appearing to be
sepals true petals lacking but outer whorl of
4-5 stamens very often petal-like, here called
petals, these frequently bilobed, and sometimes
differentiated into a long thin basal portion
(claw) and an expanded apical portion (blade or
limb) separated by appendaged joint
50Stamens 4-10, sometimes adnate to
petals Pollen tricolpate to polyporate
Carpels 2-5, connate, ovary superior, with
free-cental or occationally basal
placentation Fruit usually a loculicidal
capsule, opening by valves or apical teeth,
embryo usually curved. Floral Formula
Floral Diagram
51Distribution and Ecology Widespread, but
especially characteristic of temperate and warm
temperate regions of the Northern Hemisphere,
mostly of open habitats or disturbed
sites. Genera/species 70/2200 Major genera
Silene (700 spp.), Dianthus (300 spp.), Arenaria
(200 spp.), Gypsophila (150 spp.), Minuartia (150
spp.), Stellaria (150 spp.), Paronychia (110
spp.), and Cerastium (100 spp.). Numerous native
and introduced genera occur in the continental
United States and/or Canada. Economic plants
and products The family is best known for
ornamentals such as Dianthus (carnations, pinks),
Gypsophila (babys breath), Saponaria (soapwort),
and Silene (catchfly, campion).
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56Agrostemma githago
57Agrostemma githago
58Agrostemma githago
59Cerastium glomeratum
60Sagina nodosa
61Stellaria media
62Stellaria media
63Stellaria graminea
64Silene pratensis
65Silene regia
66Silene regia
67Silene regia
68Saponaria officinalis
69Saponaria officinalis
70Lychnis chalcedonica
71Dianthus cv. Parfait Raspberry