Berberidaceae

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Berberidaceae
W.T. Stearn (2002) The genus Epimedium and other
herbaceous Berberidaceae
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BerberidaceaeStudy day Epimedium 22-04-2006
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Position in APG II 2003 Angiosperm Phylogeny
Group(Chase et al. 2003)
Eu-Dicots
Basal eudicots
Ranunculales
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RanunculalesAPWeb(P.F. Stevens, 2001 onwards)
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Ranunculales
Ranunculaceae
Menispermaceae
Lardizabalaceae
Papaveraceae
Eupteleaceae
Berberidaceae
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Type of family Type of genus Berberis vulgaris
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CladogramBerberidaceae(molecular)chromosome-nu
mber !
(Kim Jansen 1998)
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CladogramBerberidaceae(morphological)first
clade
(Loconte Estes 1989)
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Nandina domesticawoody shrub
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Nandina domestica leaves 3-4-pinnately divided
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Nandina domesticaanthers opening with
longitudinal slits
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Nandina domestica
(Hutchinson 1973)
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CladogramBerberidaceaesecond clade fruits
1-2-seeded
(Loconte Estes 1989)
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Caulophyllum thalictroides
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Caulophyllum thalictroides staminodia
flabelliform
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Caulophyllum thalictroides fruitwall remnants,
seeds uncovered
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Gymnospermium altaicum
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Gymnospermium altaicum, G. smirnovii
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Gymnospermium shqipetarumendemic of ?
100 qindarka 1 lek
Albania
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Gymnospermium microrrhynchum
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Leontice armeniaca, L. incerta, L.
leontopetalum
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Leontice leontopetalumin the herbarium of Carl
Linnaeus
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Leontice leontopetalum
(Zohary 1966)
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Leontice incerta
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CladogramBerberidaceaethird clade stamens
tactile
(Loconte Estes 1989)
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Ranzania japonica
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Ranzania laevis Ranzania splendens
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Berberis vulgaris
(Ross-Craig 1979)
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Berberis thunbergii
(Cronquist 1981)
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Mahonia lomariifolia
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Mahonia fremontii
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Mahonia repensMahonia trifoliolata