SUMMER

1
Life Cycle of Aphthona Species Flea
Beetles Biological Control Agents of Leafy Spurge
R.D. Richard H.W. Ziolkowski, USDA-APHIS
Lana King, USDI-NPS
Adults
SUMMER
Aphthona nigriscutis
Above Ground
Aphthona lacertosa
Aphthona cyparissiae
Aphthona czwalinae
Aphthona flava
Pupae
Larvae
Below Ground
SPRING
AUTUMN
Aphthona species pupae
Aphthona larvae on leafy spurge root
Aphthona species larva
WINTER
Photos by USDA-APHIS-PPQ Bozeman Biological
Control Station
Poster by Harold Ziolkowski USDA-APHIS-PPQ