Title: Yield Practical Session 1
1FMSP stock assessment tools Training workshop
Yield Practical Session 1
2Yield Software Practical Session
- The Yield Software practical session will last
half a day. - During the session we will look in detail at
- Data requirements for the yield software.
- Then using the pre-prepared Yield Software
tutorial we will investigate some example yield
analysis with the pre-prepared example dataset.
3Loading and Saving Datasets
- To create a new dataset, open the Yield Software
and enter parameters as described in the pull
down menu then just save the parameters. - To load an existing dataset into the Yield
Software select File Load Parameters from the
menu and select the appropriate file. - To save a dataset created or modified with the
Yield Software, select File Save Parameters
from the menu and enter an appropriate filename
DO THIS OFTEN, AND WHEN HAPPY DATA ARE CORRECT.
4Entering and saving data
- First we will enter a new dataset. Open the Yield
software from the Programme menu or from the
MRAGLtd subdirectory - To create a new dataset, open the Yield Software
and enter parameters as described in the pull
down menu then just save the parameters. - We will now enter an example data set for
Lethrinus mahsena from Seychelles
5Yield Software Parameters (1/7)
- To enter or modify the parameters for the yield
software you must select the appropriate option
from the Parameters menu. Options exist for the
following sets of parameters - Von Bertalanffy
- Length-Weight
- Natural Mortality
- Maturity and Capture
- Seasonality and
- Stock-Recruit Relationship.
6Yield Software Parameters (2/7)
- Von Bertalanffy
- You will be asked to enter the parameters for the
von Bertalanffy growth curve L8, K and T0. The
growth curve will be displayed on the form. - Each parameter can be entered as a point estimate
or as a distribution. This is done by clicking
on the radio button next to the text box or the
distribution button.
7Yield Software Parameters (3/7)
- Length-Weight
- You will be asked to enter the two parameters (a
ß) for the length-weight equation - W a Lß
-
- Again these two parameters may be entered as
point estimates or as distributions.
8Yield Software Parameters (4/7)
- Natural Mortality
- You will be asked to enter a value or
distribution for the rate of natural mortality
M. - Alternatively you can enter the temperature
parameter, which (when combined with L8 and K,
that you have already entered under von
Bertalanffy) is used in Paulys equation to
estimate the natural mortality rate.
9Yield Software Parameters (5/7)
- Maturity and Capture
- Maturity and capture can be entered as ages or
lengths by toggling the button. - You will be asked to enter a value or
distribution for the length / age at maturity and
length / age at first capture. - These whether point or distribution will provide
knife-edge selection within the model.
10Yield Software Parameters (6/7)
- Seasonality of the Fishery
- You will be asked to enter the time period over
which the fishery is to be analysed (months,
years etc). - You will then be asked to indicate the breeding
and fishing seasons based on the timescale
indicated previously.
11Yield Software Parameters (7/7)
- Stock-Recruit Relationship
- You will be asked to define the stock recruit
relationship for the species. - Choices exist for three different stock-recruit
relationships, Beverton-Holt, Ricker and Constant
with their associated parameters. - You can also modify the coefficient of variation
for stochastic recruitment.
12Checking Parameters
- Built in to the software application is a
facility to cross-check parameters against each
other. - This should be run with every dataset before
starting to analyse it. - This will for instance check that the length at
first capture and maturity and less than L8, and
that the mortality rate allows fish to survive to
reach maturity. - If any parameters are not accepted, they will
need to be checked. - If all OK SAVE FILE select File Save
Parameters from the menu and enter an appropriate
filename
13Yield Example Dataset
Now we will enter the pre prepared data-set. The
yield software example dataset is based around
Lethrinus mahsena. This species is one of the
main species taken in fisheries for snappers and
emperors in the western central Indian Ocean, on
banks of the Chagos Archipelago, the Seychelles,
and Mauritius.
A data file containing estimates of parameters
and their uncertainties for L. mahsena,
LmahDat.txt, has been distributed along with the
software.
14Loading and saving data
- We will now load this existing dataset
- To load an existing dataset into the Yield
Software select File Load Parameters from the
menu and select the appropriate file. - Load LmahDat from the MRAGLtd subdirectory, take
a quick look at the data, Crosscheck parameters. - SAVE File Save Parameters from the menu and
enter an appropriate filename DO THIS OFTEN,
AND WHEN HAPPY DATA ARE CORRECT
15Number of Simulations
- The number of simulations is set by the user, by
selecting Number of simulations from the
Options menu. - Number of simulations is the number of times that
the model is run to calculate the various
different parameters the data used in each
run is sampled from the range of values
entered. It will be slightly different for each
run. - This enables us to explore the question of
uncertainty in our data - The greater the number of runs, the better, but
obviously more runs take more time.
16 Simulations, Simulating under uncertainty
17 Simulations, Simulating under uncertainty
- We will now go into Yield and look at what the
effects of both the number of runs and
statistical uncertainty in input parameters have
on the output analyses. - This is jumping ahead into analysis at this
point dont worry about interpreting the results
we look at that next. Here we are exploring
only the effects of - Number of runs
- Statistical uncertainty in input parameters
- Yield Select Equilibrium/YPR F with 100 runs
18 Simulations, Simulating under uncertainty
19 Simulations, Simulating under uncertainty
20Summary Practical Session 1
- At the end of this session you should be familiar
with
- Loading and saving datasets
- Entering and saving data
- Yield software parameters
- Checking parameters
- Simulating under uncertainty
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22FMSP stock assessment tools Training workshop
Yield Practical Session 2 Equilibrium Analyses
23Yield Software Practical Session
- The Yield Software practical session will last
half a day. - During the session we will look in detail at
- Data requirements for the yield software.
- NOW.
- Then using the pre-prepared Yield Software
tutorial we will investigate some example yield
analysis with the pre-prepared example dataset.
24Yield per Recruit Analysis (1/6)
- We will now look at the Yield-per-Recruit
Analysis - Select from the Equilibrium menu
Yield-per-Recruit vs F - This will display a dialog box as below to enter
the range and steps in F you wish to analyse.
Accept the defaults.
25Yield per Recruit Analysis (2/6)
- This will create a series of graphs against the
values of F selected showing the following - Yield per recruit
- Spawning stock biomass (SSB)
- Fishable biomass (FishB) and
- Total biomass (TotalB).
- These can be viewed as absolute values or as
fractions of the unexploited biomass.
26Yield per Recruit Analysis (3/6)
27Yield per Recruit Analysis (4/6)
- On each of the graphs the solid line shows the
mean values obtained with the confidence bands
either side. The confidence intervals are by
default set to 95 thought his can be changed by
selecting from the menu. - The data can also be viewed in a tabular format
by pressing the Medians and Intervals button. - The Input Parameters button will also show the
individual input parameters used. Note this will
not be the same for everybody!!!
28Yield per Recruit Analysis (5/6)
- What do these results mean?
- Median YPR as fraction of unexploited fishable
biomass. - Tends to a maximum at values of F gt 1.3
- Natural Mortality Rate M was based on a mean of
0.39. - Therefore F is approximately three times M, which
is quite high.
29Yield per Recruit Analysis (6/6)
- The other three biomass plots show the effect of
increasing F. - If we look at SSB-per-recruit it is clear that
levels of F producing high values of yield per
recruit are only found where the SSB is
substantially reduced in terms on its unexploited
biomass. - Note that the levels of recruitment in a YPR are
assumed constant. This is very unlikely with
such as small SSB. - Similar patterns are shown for total / fishable
biomass.
30Yield per Recruit Reference Points (1/13)
- Next we will look at the reference points that
can be calculated from Yield per recruit
analyses. - Maximum yield per recruit.
- F0.1 or F0.x
- Target spawning biomass
- Target fishable biomass
- Target total biomass
- Note that for some of the results we get a number
of impossible results.
31Yield per Recruit Reference Points (2/13)
- Select from the Equilibrium menu
Yield-per-Recruit Reference Points - This will bring up the dialog box below to allow
you to select which reference points you wish to
calculate.
32Yield per Recruit Reference Points (3/13)
- Select the defaults as before and press OK to
start calculating. - As before you will see the progress box appear.
33Yield per Recruit Reference Points (4/13)
- When complete a set of five graphs will be
displayed for the Maximum Yield per Recruit
reference point. - These show the following histograms
- Maximum yield per recruit
- Value of F that produces the maximum yield per
recruit - Spawning stock biomass per recruit
- Fishable biomass per recruit and
- Total biomass per recruit.
34Yield per Recruit Reference Points (5/13)
35Yield per Recruit Reference Points (6/13)
- The largest frequency (for F in the range 1.6
2.6) has around 30 observations. Over 50 of the
time, the F producing the maximum
yield-per-recruit exceeded 2.4. This confirms
the impression given by the yield-per-recruit
plots discussed above. - The SSB-per-recruit histogram also confirms the
impression given by the earlier plots the
largest of all the values of SSB-per-recruit was
less than 11 of its unexploited value. This is
not a good recommendation for management of a
stock.
36Yield per Recruit Reference Points (7/13)
- The maximum YPR therefore was clearly not very
useful for this stock. - Lets look at the next reference point we selected
F0.1. - Select F0.1 from the side menu by clicking on the
radio button. - Again another five graphs are displayed for this
reference point.
37Yield per Recruit Reference Points (8/13)
38Yield per Recruit Reference Points (9/13)
- Looking at the results from the analysis
presented - The most frequently occurring value of F0.1 has a
mean of 0.39, and the maximum of over 0.5. These
are obviously more sensible values of F relative
to the level of natural mortality that we
selected earlier. - What are the percentiles for F here?
- Lets display the results in a spreadsheet and
calculate them.
39Yield per Recruit Reference Points (10/13)
- As for the yield per recruit analysis itself the
data can also be viewed in a tabular format by
pressing the Medians and Intervals button. - The Input Parameters button will also show the
individual input parameters used to generate the
results shown. - Copy out the values of F from the table display
into a spreadsheet, paste into a spreadsheet and
work out the mean, median and percentiles.
40Yield per Recruit Reference Points (11/13)
- Lets look at the third reference point we
selected the target spawning biomass (set at 20
of unexploited). - Again just select the Target Spawning Biomass
button from the menu on the right hand side. - The five same graphs are displayed as for F0.1.
41Yield per Recruit Reference Points (12/13)
42Yield per Recruit Reference Points (13/13)
- The most obvious result in this form is in the
histogram for SSB-per-recruit/SSB0. As it
should, it shows that in every case, this ratio
was 20. - Looking at the histogram of values of F that
produce this, we see that most frequently, these
fell in the range 0.37 0.43, and all fell
between 0.25 and 0.79. As one would have
expected, these reference point F values are
slightly higher than those for F0.1. - Using the table of results, the median
SSB-per-recruit reference point F was 0.45 with
95 confidence interval 0.31 0.70.
43Equilibrium Yield Analysis (1/4)
- Select Equilibrium yield from the Equilibrium
menu and accept the range of F values shown. - The progress dialog box will appear while the
simulations are performed. - After a while the following form appears, and you
should select the Fraction of unexploited
biomass button.
44Equilibrium Yield Analysis (2/4)
45Equilibrium Yield Analysis (3/4)
- The main difference between yield and
yield-per-recruit methods is that for the yield
methods recruitment is allowed to vary with the
defined stock-recruit relationship. - The plot of relative yield against F suggests
that in 97.5 of the simulations, the stock was
nearly extinguished when F reached a level of 2.
- For the median, the maximum yield occurred at an
F around 0.4, and for the lower 2.5ile, F had
to lie in the range 0 0.8 to produce any
sustainable yield at all.
46Equilibrium Yield Analysis (4/4)
- The median value of the maximum yield is
estimated at around 1300t, but the percentile
range is between 600 and 2200t which is a very
large range for setting management measures. - The other plots show a similar story. In
particular, to achieve a median SSB/SSB0 ratio of
20, the corresponding F value seems to be around
0.4. - One other interesting point to note is the shape
of the plot of yield against F. The standard
Schaeffer biomass dynamic model suggests that the
yield curve is symmetric. This curve is clearly
asymmetric, with a peak shifted towards lower
values of F.
47Yield Reference Points (1/8)
- We will now look at the Yield Reference Points.
- Select Equilibrium Yield Reference Points.
- Select the default values from the menu displayed
and wait for the simulations to be carried out
and the progress dialog to be replaced by the
following set of graphs. Note that an additional
reference point has been asked for that
producing a fishable biomass at 50 of its
unexploited level.
48Yield Reference Points (2/8)
49Yield Reference Points (3/8)
- The first two graphs show that the median maximum
sustainable yield is estimated at about 1400t at
an F of 0.47. - This however seems to result in a SSB at less
than the 20 level we have assumed as a safe
level to avoid a collapse in the stock. The
fishable biomass at FMSY is at around 30 of
pre-exploitation levels, a long way below the
assumed level of the Schaefer model. (Note you
need to toggle display option to Fraction of
unexploited biomass to see these) - Lets now look at the Target Spawning Biomass
reference point.
50Yield Reference Points (4/8)
51Yield Reference Points (5/8)
- The 20 target spawning biomass reference point
gives a reasonable value for F of approximately
0.43, which is within reasonable bounds compared
to M. - Check the confidence intervals for F in the
results table. - The final reference point to look at is the 50
fishable biomass reference point.
52Yield Reference Points (6/8)
53Yield Reference Points (7/8)
- Again, we would expect this to be achieved at a
lower value of F, given that the median fishable
biomass ratio producing MSY was estimated earlier
to be 30. - Now, the mode occurs at an F around 0.211. Using
the table of results, the median was estimated to
be 0.20, with 95 confidence interval 0.15
0.26. - All these analyses are heavily dependent on the
input parameters and we should test the
sensitivity of these.
54Yield Reference Points (8/8)
- One of the parameters that is more doubtful than
others is the R0 parameter (number of recruits
before exploitation) of the stock recruit
relationship. - This was set at 25 million initially.
- Try values of 2.5, 10 and 250 million and
investigate the results. Are the reference
points sensitive to the R0 parameter?
55Summary Practical Session 2
- Equilibrium models
- Yield-per-recruit analyses (constant recruitment)
- Yield analyses (deterministic recruitment)
- Biological Reference Points (BRPs)
- Yield (MSY)
- Biomass (SSB20, Fishable Biomass, Total Biomass)
- Fishing Effort (F0.1)
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57FMSP stock assessment tools Training workshop
Yield Practical Session 3 Transient Analyses
58Transient Analysis (1/6)
- We have assumed so far that the population has
been in equilibrium. - Let us calculate the transient SSB reference
point. - We will look for a probability (10) that the
stock will remain above a proportion of SSB (20)
for twenty years. - Select the Transient SSB Reference Point from the
Transient menu and accept the default values as
above.
59Transient Analysis (2/6)
- This will then perform the simulation giving a
probability estimate based on our parameters. - Here for instance we have calculated that an F of
0.271 would give a probability that the
population will drop below 20 in one or more
years in the next twenty.
60Transient Analysis (3/6)
- This suggests that, with even relatively modest
amounts of recruitment variability the risks of
the SSB falling below specified low levels can be
rather greater than might have been imagined. - The value we have obtained is rather less than
the median FMSY (0.41), but then we would expect
that because that F on average reduces the SSB to
less than 20 of its unexploited level even when
there is no recruitment variability. With
variability there is always the possibility of a
number of bad recruitment years occurring one
after the other.
61Transient Analysis (4/6)
- We can now look at projecting a strategy for
fisheries management into the future on our
imaginary stock. - Select Transient Projections from the menu.
- Then project the stock into the future by using
the value of F you obtained in the Transient SSB
Reference Point. We assume that this will on
average allow the SSB to drop below 20 of SSB0
in 10 of the years. - What do you find?
62Transient Analysis (5/6)
- Try the same analysis for the values of F for the
FMSY and the SSB/SSB0 20 target reference points
you obtained. - What happens to the stock?
- Try a few other scenarios to see what happens,
e.g. unrestricted fishing at very high values of
F or periods of no fishing between periods of
high fishing.
63Summary Reference Points
Estimated Fishing Mortality Rates (F) from
Equilibrium and Transient Analyses when Lc 22.8
cm
64Summary - Managing under uncertainty
- Managers base their decisions on certain
reference points that indicate a particular
course of action. Yield derives these. However we
have seen that statistical uncertainty in the
biological data estimated for a fishery, and
inter-annual variation in the SRR can lead to
considerable uncertainty in the values of the
reference points and thus in management
decisions. - Yield quantifies this uncertainty and enables
managers to make more informed decisions about
how best to manage. - Yield can be a powerful tool when used properly
and in conjunction with other stock assessment
tools.